Fig. 1

Simulated animal movement tracks (300 steps per track) on a randomly generated landscape displaying behaviours consistent with each hypothesis (and model). The colour of each point on this simulated movement track represents the hypothetical time in the animal’s memory “cycle”, which is here set to 100 time units (points at \(t = 75\) have the same colour as \(t = 175\)). The null model implies completely random movement, while the resource-only model implies that the animal will locate nearby resources and select for those areas. The memory-only model implies that the animal relocates itself to areas it visited 100 time units before. The resource-memory model combines mechanisms in the resource-only and memory-only models