Per reef patch
| |
I. Local settlement
|
LRa
|
Local retention
|
SR
|
Self-recruitment (with eight-patch metapopulation)
|
H’
|
Shannon index of diversity of settlers (sensitive to weak connections)
|
II. Downstream connectivity
|
mdGa
|
Median geographic distance of downstream connections
|
mxG
|
Maximum distance of downstream connections
|
S
|
Total proportion of larvae that settle downstream
|
dC
|
Out-degree, total number of downstream connections
|
Cw
a
|
Weighted degree centrality as dC(1-α) x Sα; dp is d as proportion of total possible connections; α = 0.5
|
III. Metapopulation consequences
|
λM
a
|
Metapopulation growth rate with variable population sizes, fecundity, & survival [6]
|
λmax
|
Metapopulation capacity [5]
|