Per reef patch
 
I. Local settlement

LR^{a}

Local retention

SR

Selfrecruitment (with eightpatch metapopulation)

H’

Shannon index of diversity of settlers (sensitive to weak connections)

II. Downstream connectivity

mdG^{a}

Median geographic distance of downstream connections

mxG

Maximum distance of downstream connections

S

Total proportion of larvae that settle downstream

dC

Outdegree, total number of downstream connections

C_{w}
^{a}

Weighted degree centrality as dC^{(1α)} x S^{α}; dp is d as proportion of total possible connections; α = 0.5

III. Metapopulation consequences

λ_{M}
^{a}

Metapopulation growth rate with variable population sizes, fecundity, & survival [6]

λ_{max}

Metapopulation capacity [5]
